Three men were charged by police over the incidents at White Hart Lane, which caused a 10-minute suspension of the game late in the first half.
UEFA’s control, ethics and disciplinary body (CEDB) announced the fine, while Nathaniel Thompson, 23, Helder Gomes, 22, and Daniel Jarvis, 26, were arrested after encroaching onto the pitch.
All three were wearing branded T-shirts, reportedly of headphone manufacturer BassBuds, and the stunt appeared to be orchestrated as they tried to take photos with the Spurs players.
BassBuds company, which is associated with Spurs and six other Premier League teams, issued a statement via Facebook condemning the actions of three rogue pranksters during the filming of an online hidden camera show.
Match referee Yevhen Aranovskiy had warned the tie on 27 November, which Spurs won 1-0, would be cancelled were a fourth spectator to invade the pitch.
-express